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Blackwatch, a Fort Detrick special forces unit dedicated to combating biological warfare, and the Infected, consisting of civilians who have contracted a strange virus and powerful monsters known as Hunters. During his quest, Alex faces two factions. With no memory of his previous life, Alex is forced to track down and consume those related to the conspiracy in order to uncover the truth.

He soon discovers that he now possesses powerful shapeshifting abilities that grant him superhuman strength, speed, agility, durability, weaponry and the ability to " consume" people to gain their memories, skills and appearance. Alex escapes and sets out to recover his memory and find out exactly what occurred. Mercer (voiced by Barry Pepper) who wakes up on a morgue slab in the basement of Gentek, a large and influential genetic engineering company.

The game follows the story of Alexander J.